Uca intermedia

Following is a fairly comprehensive list of papers, books, and theses that deal or refer to fiddler crabs. The list currently contains 2,486 references (last updated October 17, 2007). Many of these papers (particularly the older ones) are primarily taxonomic lists. I tried to leave out papers which just casually refer to fiddlers without providing any information (e.g., “...the results of these experiments on the lobster are similar to those seen in green crabs and fiddler crabs.”). In a list of this size, there are bound to be errors, omissions, and mistaken inclusions, but I suspect you'll still find it a pretty good reference. Feel free to send me corrections.

The list is also available as a zipped Endnote file (Version 10 for Windows).
[Note: may not be compatible with older versions]


1983. Recent traces of marine endofauna from Qingdau. Earth Science (Wuhan) 20:114.

Abbott, J. F. 1913. The effect of distilled water upon the fiddler crab. Biological Bulletin 24(3):169-174.

Abbott, W., and C. E. Dawson. 1966. Response of some anomuran and brachyuran crustaceans to high-speed centrifugation. Nature 211:1320-1321.

Abele, L. G. 1972. Comparative habitat diversity and faunal relationships between the Pacific and Caribbean Decapod Crustacea of Panama. Pp. 114 (Coral Gables, Florida: University of Miami).

Abele, L. G., and W. Kim. 1986. An Illustrated Guide to the Marine Decapod Crustaceans of Florida. Tallahassee, FL: State of Florida Department of Environmental Regulation.

Abele, L. G., and W. Kim. 1989. The Decapod Crustaceans of the Panama Canal. Smithsonian Contributions to Zoology 482:1-50.

Abramowitz, A. A. 1937. The chromatophorotropic hormone of the Crustacea: Standardization, properties and physiology of the eye-stalk glands. Biological Bulletin 72(3):344-365.

Abramowitz, A. A., and R. K. Abramowitz. 1938. On the specificity and related properties of the crustacean chromatophorotropic hormone. Biological Bulletin 74(2):278-296.

Abramowitz, R. K., and A. A. Abramowitz. 1940. Moutling, growth, and survival after eyestalk removal in Uca pugilator. Biological Bulletin 78(2):179-188.

Abramson, C. I., and R. D. Feinman. 1986. Discriminant and yoked punishment in the fiddler crab Uca pugilator. Society for Neuroscience Abstracts 12(1):40.

Abreu, V. 1975. Fiddlers on the shorelines. Of Sea & Shore 6(3):139-140.

Adamowicz, S. J., and A. Purvis. 2006. From more to fewer? Testing an allegedly pervasive trend in the evolution of morphological structure. Evolution 60(7):1402-1416.

Adams, A., and A. White. 1848. Crustacea. Pp. 1-66 in The zoology of the voyage of H.M.S. "Samarang"; under the command of Captain Sir Edward Belcher, during the years 1843-1846, A. Adams, ed., Volume 8. London: Reeve, Benham, & Reeve.

Adirwiryono, S., Sukristiyono, and V. Toro. 1984. The occurrence of crustaceans in the Tanjung Bungin mangrove forest, South Sumatara, Indonesia. Pp. 241-257 in Proceedings of the Asian Symposium on Mangrove Envrionmental Research and Management held in Kuala Lumpur, 25-29 August 1980, E. Soepadmo, A. N. Rao and D. J. Macintosh, eds. Kuala Lumpur: Univeristy of Malaya & Unesco.

Ahmed, M. 1976. A study of the normal and abberrant sexual types of the Venezuelan fiddler crabs Uca cumulanta and U. rapax. Bulletin of Marine Science 26(4):499-505.

Ahmed, M. 1978. Development of asymmetry in the fiddler crab Uca cumulanta Crane, 1943 (Decapoda Brachyura). Crustaceana 34(3):294-300.

Ahmed, M., and R. A. Kahn. 1976. The occurrence of larval trematode in populations of the fiddler crab Uca lactea (De Haan, 1835) (Decapoda, Ocypodidae) on the coast of Karachi, Pakistan. Crustaceana 31(2):216-217.

Ahmed, M., and R. A. Khan. 1978. Development of asymmetry in the claws of the fiddler crab Uca lactea. Pakistan Journal of Zoology 10(1):49-54.

Aicher, B. 1987. Vibratorische Kommunikation bei der Winkerkrabbe Uca pugilator (Brachyura, Ocypodidae): Vibrationssignale, ihre substratgebundene Fortleitung und neuronale Rezeption: Konstanz).

Aicher, B., H. Markl, W. M. Masters, and H. L. Kirschenlohr. 1983. Vibration transmission through the walking legs of the fiddler crab, Uca pugilator (Brachyura, Ocypodidae) as measured by laser doppler vibrometry. Journal of Comparative Physiology 150A(4):483-491.

Aicher, B., and J. Tautz. 1984. 'Peripheral inhibition' of vibration-sensitive units in the leg of the fiddler crab Uca pugilator. Journal of Comparative Physiology 154A(1):49-52.

Aicher, B., and J. Tautz. 1990. Vibrational communication in the fiddler crab, Uca pugilator. I. Signal transmission through the substratum. Journal of Comparative Physiology 166A(3):345-353.

Aikawa, H. 1937. Further notes on brachyuran larvae. Records of Oceanographic Works in Japan 9:87-162.

Aizawa, N. 2000. Heterospecific timing of waving display in ocypodid crabs Ilyoplax pusilla and Uca lactea. Journal of the Marine Biological Association of the United Kingdom 80(5):947-948.

Albrecht, H., and H.-O. von Hagen. 1981. Differential weighting of electrophoretic data in crayfish and fiddler crabs (Decapoda: Astacidae and Ocypodidae). Comparative Biochemistry and Physiology 70B:393-399.

Alcock, A. 1892. On the habits of Gelasimus annulipes, Edw. Annals and Magazine of Natural History, Series XI 10:415-416.

Alcock, A. 1900. Materials for a carcinological fauna of India No. 6. The Brachyura Catometopa, or Grapsoidea. Journal of the Asiatic Society of Bengal 64(2):279-486.

Alcock, A. 1902. A Naturalist in Indian Seas. London.

Alcock, A., and A. R. Anderson. 1894. Natural history notes from H.M. Indian marine survey steamer "Investigator," Commander C. F. Oldham, R.N., commanding. Series 11, No. 14. An account of a recent collection of deep-sea Crustacea from the Bay of Bengal, Laccadive Sea. Journal of the Asiatic Society of Bengal 63:141-185.

Alcock, J. J., Jr. 1997. Hood-Building in the Fiddler Crab (Uca musica musica)in Neurobiology and Behavior (Ithaca, NY: Cornell University).

Allen, B. J., and J. S. Levinton. 2002. Fiddler crab fitness: Exercise and the cost of sex. Integrative and Comparative Biology 42(6):1184.

Allen, B. J., and J. S. Levinton. 2005. The paradox of the weakening combatant: Trade-off between closing force and gripping speed in a sexually selected combat structure. Integrative and Comparative Biology 44(6):?

Allen, B. J., and J. S. Levinton. 2007. Costs of bearing a sexually selected ornamental weapon in a fiddler crab. Functional Ecology 21(1):154-161.

Allen, D. M., W. S. Johnson, and V. Ogburn-Matthews. 1995. Trophic relationships and seasonal utilization of salt-marsh creeks by zooplanktivorous fishes. Environmental Biology of Fishes 42(1):37-50.

Allen, E. A., and H. A. Curran. 1973. Lebensspuren of selected decapod crustaceans in recent lagoon margin and estuarine environments. Abstracts with Programs 4(1):1.

Allen, E. A., and H. A. Curran. 1974. Biogenic sedimentary structures produced by crabs in lagoon margin and salt marsh environments near Beaufort, North Carolina. Journal of Sedimentary Petrology 44(2):538-548.

Allen, R. K. 1976. Common Intertidal Invertebrates of Southern California. Revised Edition Palo Alto, CA: Peek Publications.

al-Mossawi, L. S. M. E. 2001. Seasonal and lunar variation in the tidal rhythm of emergence of the fiddler crab Uca lactae annuilepus, at a shore in Kuwait. Chronobiology International 18(6):1095-1096.

Altevogt, R. 1955. Beobachtungen und Untersuchungen an indischen Winkerkrabben. Zeitschrift für Morphologie und Ökologie der Tiere 43(6):501-522.

Altevogt, R. 1955. Some studies on two species of Indian fiddler crabs, Uca marionis nitidus (Dana) and U. annulipes (Latr.). Journal of the Bombay Natural History Society 52:702-716.

Altevogt, R. 1956. Der Mechanismus der Nahrungsaufnahme bei Winkerkrabben. Naturwissenschaften 43(4):92-93.

Altevogt, R. 1956. Neue Untersuchungen an indischen Winkerkrabben. Pp. 148-150 in Verhandlungen der Deutschen Zoologischen Gesellschaft in Hamburg. Leipzig: Akademische Verlagsgesellschaft.

Altevogt, R. 1957. Beiträge zur Biologie und Ethologie von Dotilla blanfordi Alcock und Dotilla myctiroides (Milne Edwards) (Crustacea Decapoda). Zeitschrift für Morphologie und Ökologie der Tiere 46:369-388.

Altevogt, R. 1957. Untersuchungen zur Biologie, Ökologie und Physiologie Indischer Winkerkrabben. Zeitschrift für Morphologie und Ökologie der Tiere 46(1):1-110.

Altevogt, R. 1959. The love and life of fiddler crabs. Journal of the Bombay Natural History Society 56:1-7.

Altevogt, R. 1959. Ökologische und ethologische Studien an Europas einziger Winkerkrabbe Uca tangeri Eydoux. Zeitschrift für Morphologie und Ökologie der Tiere 48(2):123-146.

Altevogt, R. 1959. Winkerkrabben. Kosmos, Stuttgart 55:60-66.

Altevogt, R. 1959. Zur Ökologie und Ethologie von Uca tangeri (Eydoux), Europas einziger Winkerkrabbe. Proceedings of the International Congress of Zoology 15:891-892.

Altevogt, R. 1962. Akustische Epiphänomene im Sozialverhalten von Uca tangeri in Südspanien. Verhandlungen der Deutschen Zoologischen Gesellschaft 26:309-315. {Acoustical epiphenomena in the social behavior of Uca tangeri in southern Spain}

Altevogt, R. 1963. Lernversuche bei Uca tangeri. Zoologische Beiträge 9:447-459.

Altevogt, R. 1963. Wirksamkeit polarisierten Lichtes bei Uca tangeri. Naturwissenschaften 50(22):697-698.

Altevogt, R. 1964. Ein antiphoner Klopfkode und eine neue Winkfunktion bei Uca tangeri. Naturwissenschaften 51:644-645.

Altevogt, R. 1965. Lichtkompass- und Landmarkendressuren bei Uca tangeri in Andalusien. Zeitschrift für Morphologie und Ökologie der Tiere 55(6):641-655.

Altevogt, R. 1965. Uca tangeri (Eydoux, 1835) in der terra typica. Crustaceana 8:31-36.

Altevogt, R. 1968. The foam bath of brachyuran crustaceans as temperature regulator. Zoologischer Anzeiger 181(5-6):398-402.

Altevogt, R. 1969. Das "Schaumbaden" brachyurer Crustaceen als Temperatur-Regulator. Zoologischer Anzeiger 181:5-6.

Altevogt, R. 1969. Ein sexualethologischer Isolationsmechanismus bei sympatrischen Uca-Arten (Ocypodidae) des Ostpazifik. forma et functio 1:238-249. {An ethological reproductive isolation mechanism in sympatric species of Uca (Ocypodidae) of the Eastern Pacific}

Altevogt, R. 1970. Form und Funktion der vibratorischen Signale von Uca tangeri und Uca inaequalis (Crustacea, Ocypodidae). forma et functio 2:178-187. {Form and function of vibrational signals in Uca tangeri and Uca inaequalis (Crustacea, Ocypodidae)}

Altevogt, R. 1972. Physiological inter-relations of display and locomotion in fiddler crabs: An evolutionary aspect. Journal of the Marine Biological Association of India 14(2):456-467.

Altevogt, R. 1978. Proto-fiddlers and fiddlers: Pathways to waving in Indian Brachyuran crabs. Journal of the Bombay Natural History Society 75(3):800-809.

Altevogt, R. 1988. Fiddlers on the retreat? Pp. 367-370 in Proceedings of the Symposium on Endangered Marine Animals and Marine Parks. Cochin, India 12-16 January, 1985, E. G. Silas, ed. Cochin, India: Marine Biological Association of India.

Altevogt, R., and T. A. Davis. 1979. Shift of handedness in Indian populations of Uca vocans (Linné 1758) (Crustacea, Decapoda, Ocypodidae). Zoologischer Anzeiger 203(5-6):342-344.

Altevogt, R., and H.-O. von Hagen. 1964. Über die Orientierung von Uca tangeri Eydoux im Freiland. Zeitschrift für Morphologie und Ökologie der Tiere 53:636-656.

Alvarez Del Castillo-C, M., M. E. Hendrickx, and S. Rodriguez-C. 1992. Decapod crustaceans from the Barra de Navidad Lagoon, Jalisco, Mexico. Proceedings of the San Diego Society of Natural History 27:1-9.

Alvarez, R. Z. 1968. Crustáceos Decápodos Ibéricos. Investigacíon Pesquera (Barcelona) 32:1-510.

Anastasia, J. R., S. G. Morgan, and N. S. Fisher. 1998. Tagging crustacean larvae: Assimilation and retention of trace elements. Limnology and Oceanography 43(2):362-368.

Andersen, M. L., M. Bohn, J. T. Høeg, and P. G. Jensen. 1990. Cyprid ultrastructure and adult morphology in Ptychascus barnwelli, new species, and P. glaber (Cirripedia: Rhizocephala), parasites on semiterrestrial crabs. Journal of Crustacean Biology 10(1):20-28.

Andrews, P. M. 1973. Ultrastructural study of the pericardial organ-anterior ramifications complex neurosecretory terminals. Zeitschrift für Zellforschung und mikroskopische Anatomie 144(3):309-324.

Anger, K., M. Montú, C. de Bakker, and L. Loureiro Fernandes. 1990. Larval development of Uca thayeri Rathbun, 1900 (Decapoda: Ocypodidae) reared in the laboratory. Meeresforschung 32(4):276-294.

Anger, K., E. D. Spivak, C. Bas, D. Ismael, and T. Luppi. 1994. Hatching rhythms and dispersion of decapod crustacean larvae in a brackish coastal lagoon in Argentina. Helgolaender Meeresuntersuchungen 48(4):445-466.

Anilkumar, G., P. M. Hopkins, and D. S. Durica. 2006. Comparative studies of the EcR and RXR genes in allopatric Uca species. Integrative and Comparative Biology 46(6):?

Aoki, H., M. N. Ahsan, K. Matsuo, T. Hagiwara, and S. Watabe. 2003. Purification and characterization of collagenolytic proteases from the hepatopancreas of Northern shrimp (Pandalus eous). Journal of Agricultural and Food Chemistry 51(3):777-783.

Armendariz, L. C. 2005. Redescription of the megalopa of the fiddler crab Uca uruguayensis (Decapoda, Brachyura, Ocypodidae) with special emphasis on its setae. Journal of Natural History 39(22):1981-1993.

Armendariz, L. C., and I. I. Cesar. 2006. Estimating secondary production of Uca uruguayensis (Decapoda, Brachyura, Ocypodidae) from Samborombon Bay, Argentina. Ciencias Marinas 32(3):551-558.

Arnoux, B., A. Ducruix, and A. Lecroisey. 1990. Modélisation de la Protéase I extraite du crabe Uca pugilator. Pathologie Biologie 38(10):975-981. {A model of protease I from the fiddler crab Uca pugilator}

Arnoux, B., A. Lecroisey, and A. Ducruix. 1990. Modeling of protease I collagenolytic enzyme from the fiddler crab Uca pugilator. Journal of Computer-Aided Molecular Design 4(2):107-116.

Arnqvist, G. 2006. Sensory exploitation and sexual conflict. Philosophical Transactions of the Royal Society 361B(1466):375-386.

Ashton, E. C., P. J. Hogarth, and D. J. Macintosh. 2003. A comparison of brachyuran crab community structure at four mangrove locations under different management systems along the Melaka Straits-Andaman Sea Coast of Malaysia and Thailand. Estuaries 26(6):1461-1471.

Ashton, E. C., D. J. Macintosh, and P. J. Hogarth. 2003. A baseline study of the diversity and community ecology of crab and molluscan macrofauna in the Sematan mangrove forest, Sarawak, Malaysia. Journal of Tropical Ecology 19:127-142.

Aspey, W. P. 1971. Inter-species sexual discrimination and approach-avoidance confict in two species of fiddler crabs, Uca pugnax and Uca pugilator. Animal Behaviour 19(4):669-676.

Aspey, W. P. 1978. Fiddler crab behavioral ecology: Burrow density in Uca pugnax (Smith) and Uca pugilator (Bosc) (Decapoda Brachyura). Crustaceana 34(3):235-244.

Atkinson, R. J. A., and A. C. Taylor. 1988. Physiological ecology of burrowing decapods. Pp. 201-226 in Aspects of Decapod Crustacean Biology, A. A. Fincham and P. S. Rainbow, eds. Oxford: Clarendon Press.

Audouin, M. M. V., and H. Milne Edwards. 1829. Histoire naturelle des annelides, crustaces et arachnides. Pp. 1-252 in Resume d'Entomologie, ou d'Histoire Naturelle des Animaux Articules, Volume I. Paris: Bureau de l'Encyclopedie Portative.

Aurivillius, C. W. S. 1893. Die Beziehungen der Sinnesorgane amphibischer Decapoden zur Lebensweise und Athmung. Nova Acta R. Soc. Scient. Upsal. Nyt. Mag. Naturvid 3:1-48.

Aurivillius, C. W. S. 1898. Krustaceen aus dem Kamerun. Gebiete. Bih. K. svenska Vetensk Akad. Handl. 24 afd. 4(1):1-31.

Avent, R. M., Jr. 1973. The effects of hydrostatic pressure and temperature on Uca pugilator (Bosc). Pp. 97: Florida State University).

Avent, R. M., Jr. 1974. The effects of hydrostatic pressure on living aquatic organisms. VIII. Behavioral and metabolic responses of Uca pugilator to variations in hydrostatic pressure and temperature. Internationale Revue der gesamten Hydrobiologie 59(2):219-238.

Avent, R. M., Jr. 1975. Evidence for acclimation to hydrostatic pressure in Uca pugilator (Crustacea: Decapoda: Ocypodidae). Marine Biology 31(2):193-199.

Azzalina, J. D., and D. G. Trainer. 1985. Amylolytic activity in the hepatopancreas of Uca minax, Uca pugnax and Uca pugilator. Comparative Biochemistry and Physiology 82B(4):679-682.

Bachand, R. G. 1979. The fascinating fiddler crab. Underwater Naturalist 11(4):12-14.

Bachmann, K., and E. L. Rheinsmith. 1973. Nuclear DNA amounts in Pacific Crustacea. Chromosoma 43(3):225-236.

Backwell, P. R. Y., J. H. Christy, S. R. Telford, M. D. Jennions, and N. I. Passmore. 2000. Dishonest signalling in a fiddler crab. Proceedings of the Royal Society of London 267B(1444):719-724.

Backwell, P. R. Y., and M. D. Jennions. 2004. Coalition among male fiddler crabs. Nature 430(6998):417.

Backwell, P. R. Y., M. D. Jennions, J. H. Christy, and N. I. Passmore. 1999. Female choice in the synchronously waving fiddler crab Uca annulipes. Ethology 105(5):415-421.

Backwell, P. R. Y., M. D. Jennions, J. H. Christy, and U. M. Schober. 1995. Pillar building in the fiddler crab Uca beebei: Evidence for a condition-dependent ornament. Behavioral Ecology and Sociobiology 36:185-192.

Backwell, P. R. Y., M. D. Jennions, N. I. Passmore, and J. H. Christy. 1998. Synchronized courtship in fiddler crabs. Nature 391:31-32.

Backwell, P. R. Y., M. D. Jennions, K. Wada, M. Murai, and J. H. Christy. 2006. Synchronous waving in two species of fiddler crabs. Acta Ethologica 9(1):22-25.

Backwell, P. R. Y., P. D. O'Hara, and J. H. Christy. 1998. Prey availability and selective foraging in shorebirds. Animal Behaviour 55(6):1659-1667.

Backwell, P. R. Y., and N. I. Passmore. 1996. Time constraints and multiple choice criteria in the sampling behaviour and mate choice of the fiddler crab, Uca annulipes. Behavioral Ecology and Sociobiology 38(6):407-416.

Bairagi, N., and A. Misra. 1988. On the taxonomic status of Gelasimus acutus Stimpson (Decapoda: Ocypodidae) present in the national collection of the Zoological Survey of India, Calcutta. Journal of the Bombay Natural History Society 85(2):449-451.

Baird, J. L., Jr., and A. L. Burleson. 1970. An analysis of locomotor behavior in the fiddler crab Uca pugilator. American Zoologist 10(4):500.

Balbort, L., and C. L. Thurman, II. 1997. Comparison of locomotor rhythms in sympatric Uca from the western Gulf of Mexico. American Zoologist 37(5):186A.

Baldwin, G. F. 1976. Hyper- and hypo-tonic regulation in the fiddler crab, Uca spp. Pp. 70: Washington State University).

Baldwin, G. F., and L. B. Kirschener. 1976. Sodium and chloride regulation in Uca adapted to 10% sea water. Physiological Zoölogy 49(2):172-180.

Baldwin, G. F., and L. B. Kirschener. 1976. Sodium and chloride regulation in Uca adapted to 175% sea water. Physiological Zoölogy 49(2):158-171.

Balss, H. 1921. Über Stridulations Organe bei dekapoden Crustaceen. Eine zusammenfassende Übersicht. Naturw. Wochenschr. Jena, neue Folge 20:697-701.

Balss, H. 1922. Crustacea VII; Decapoda Brachyura (Oxyrhyncha und Brachyrhyncha) und geographische Übersicht über Crustacea Decapoda. Pp. 71-110 in Beitr. Kennt. Meeresfauna Westafr., H. v. W. Michaelson, ed., Volume 3. Hamburg

Balss, H. 1922. Ostasiatische Decapoden. IV. Die Brachyrhynthen (Cancridea). Archiv für Naturgeschicthe 88A(11):94-166.

Balss, H. 1924. Expedition S. M. Schiff "Pola" in das Rote Meer 1895/6-1897/8. Zool. Ergeb. XXXIV. Decapoden des Roten Meeres. III. Die Parthenopiden. Cylco- und Catometopen. Denkschr. Akad. Wiss. Wien 99:1-18.

Balss, H. 1938. Die Dekapoda Brachyura von Dr. Sixten Bocks' Pazifik-Expedition, 1917-1918. Göteborgs Kungliga Vetenskaps-och Vitterhetssam-halles handlinger (5B) 5(7):1-85.

Balss, H. 1955. Decapoda. Brons Klassen 5(Abt. 1 Buch 7):1285-1367.

Barden, H., and S. Koulish. 1983. The dark brown integumentary pigment of a barnacle (Balanus eburneus). A comparative study. Histochemistry 78(1):41-52.

Barnard, K. H. 1950. Descriptive catalogue of South African decapod Crustacea. Annals of the South African Museum 38:1-837.

Barnes, W. J. P. 1975. Leg co-ordination during walking in the crab, Uca pugnax. Journal of Comparative Physiology 96:237-256.

Barnes, W. J. P. 1975. Nervous control of locomotion in Crustacea. Pp. 415-441 in 'Simple' Nervous Systems, P. N. R. Underwood and D. R. Newth, eds. London: Arnold.

Barnwell, F. H. 1963. Observations on daily and tidal rhythms in some fiddler crabs from Equatorial Brazil. Biological Bulletin 125(3):399-415.

Barnwell, F. H. 1965. Comparative studies on daily and tidal rhythms in fiddler crabs (genus Uca) from the Woods Hole region. Pp. 122 (Evanston, Illinois: Northwestern Univsersity).

Barnwell, F. H. 1966. Daily and tidal patterns of activity in individual fiddler crab (genus Uca) from the Woods Hole region. Biological Bulletin 130(1):1-17.

Barnwell, F. H. 1968. Comparative aspects of the chromatophoric responses to light and temperature in fiddler crabs of the genus Uca. Biological Bulletin 134(2):221-234.

Barnwell, F. H. 1968. The role of rhythmic systems in the adaptation of fiddler crabs to the intertidal zone. American Zoologist 8(3):569-583.

Barnwell, F. H. 1976. Variation in the form of the tide and some problems it poses for biological timing systems. Pp. 161-187 in Biological Rhythms in the Marine Environment, P. J. DeCoursey, ed., Volume 4. Columbia, SC: University of South Carolina Press.

Barnwell, F. H. 1980. Taxonomic differentiation within the lactea-complex of fiddler crabs in the Sunda Shelf region. American Zoologist 20:957.

Barnwell, F. H. 1982. The prevalence of male right-handedness in the Indo-West Pacific fiddler crabs Uca vocans (Linnaeus) and U. tetrogonon (Herbst) (Decapoda: Ocypodidae). Journal of Crustacean Biology 2(1):70-83.

Barnwell, F. H. 1984. Tidal rhythms of locomotor activity in the fiddler crab Uca vocans from shores with semi-diurnal and predominantly diurnal. American Zoologist 24(3):68A.

Barnwell, F. H. 1986. Fiddler crabs of Jamaica (Decapoda, Brachyura, Ocypodidae, genus Uca). Crustaceana 50(2):146-165.

Barnwell, F. H. 1986. The functional morphology of external genitalia in fiddler crabs genus Uca. American Zoologist 26(4):34A.

Barnwell, F. H. 1988. Stereo-imaging of external genitalia in the neotropical vocator-complex of fiddler crabs. American Zoologist 28(4):133A.

Barnwell, F. H. 2002. Daily and tidal rhythms of locomotor activity in the fiddler crab Uca tetragonon in relation to the "solar" tide at Moorea, French Polynesia. Integrative and Comparative Biology 42(6):1191.

Barnwell, F. H., and F. A. Brown, Jr. 1964. Differences in the persistent metabolic rhythms of fiddler crabs from two levels of the same beach. Biological Bulletin 125(2):371-372.

Barnwell, F. H., and J. Martini. 1993. Daily and tidal rhythms of activity in the Andalusian fiddler crab, Uca tangeri. American Zoologist 33(5):91A.

Barnwell, F. H., and J. A. Matoyoshi. 1970. Protein differences in new-world fiddler crabs Brachyura genus Uca. American Zoologist 10(4):505.

Barnwell, F. H., K. Shankaraiah, and F. Halberg. 1983. Linear nonlinear rhythmometry of activity in fiddler crabs Uca princeps from a semi diurnal tidal habitat. Chronobiologia 10(2):109.

Barnwell, F. H., and J. H. Stillman. 1990. Differences in daily and tidal rhythms in populations of the fiddler crabs Uca princeps and U. monilifera. American Zoologist 30(4):138A.

Barnwell, F. H., and W. A. Szelistowski. 1985. Twenty-one species of fiddler crabs Uca from a small tidal river on the Pacific coast of Costa-Rica. American Zoologist 25(4):86A.

Barnwell, F. H., and C. L. Thurman, II. 1983. Endemism and faunal transition areas for fiddler crabs Uca spp. in the Gulf of Mexico. American Zoologist 23(4):947.

Barnwell, F. H., and C. L. Thurman, II. 1984. Taxonomy and biogeography of fiddler crabs (Ocypodidae: genus Uca) of the Atlantic and Gulf coasts of eastern North America. Zoological Journal of the Linnean Society 81:23-87.

Barnwell, F. H., M. C. Vanzwol, and R. C. Zinnel. 1987. Geographic variation in allometry of the buccal cavity in the fiddler crabs Uca pugilator and Uca panacea. American Zoologist 27(4):141A.

Barnwell, F. H., and K. C. Zinnel. 1981. The fiddler crab Uca spp. a model for human multi-oscillator circadian organization. International Journal of Chronobiology 7(4):11.

Barnwell, F. H., and K. C. Zinnel. 1984. The fiddler crab: A model for the study of internal desynchronization of circadian rhythms. Pp. 250-256 in Chronobiology 1982-1983, E. Haus and H. F. Kabat, eds. Basel: S. Karger.

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