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Al-Ghais & Cooper (1996)

Al-Ghais, S.M., and R.T. Cooper (1996) Brachyura (Grapsidae, Ocypodidae, Portunidae, Xanthidae and Leucosiidae) of Umm Al Quwain mangal, United Arab Emirates. Tropical Zoology 9:409–430.

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Naderloo et al. (2010)