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Species: Ocypoda pugilator
Subgenus: Minuca
Common Name: Atlantic Sand Fiddler Crab
Geographic Range (Map): Western Atlantic
- USA (Massachusetts to Mississippi, scattered in Texas), Bahamas
This drawing is from De Kay (1844). He called this species Gelasimus vocans. It is almost certainly Uca pugilator. A second figure of a "Variety A" is probably Uca pugnax. Note that the major claw is drawn backwards (i.e., it is a right major claw on a left-handed crab). This figure was probably based on a collected specimen missing the major cheliped and the one they replaced it with for the drawing was a right (and not a left) major cheliped.
This photo was taken from a website at Francis Marion University.
From Tarpon Springs Notebook. I'm pretty sure these are U. pugilator.
From an Australian web site.
Male and female on rocky background
Photos by M. S. Rosenberg, Flax Pond, Long Island, New York.
This photo is from www.webshots.com.
These photos are from Petwer Woods photography.
These photos are from Florida and were provided by Chris Lukhaup.
Eight week old crabs. The photos were provided by Hans-Georg Rupp.
No video available at this time.
Abele and Kim (1986); Abramowitz and Abramowitz (1940); Abramson and Feinman (1986); Aicher (1987); Aicher and Tautz (1984, 1990); Aicher et al. (1983); Allen and Levinton (2002); Altevogt (1957, 1959); Anilkumar et al. (2006); Aoki et al. (2003); Arnoux et al. (1990 x 2); Aspey (1971, 1978); Atkinson and Taylor (1988); Avent (1973, 1974, 1975); Azzalina and Trainer (1985); Bachmann and Rheinsmith (1973); Baird and Burleson (1970); Barden and Koulish (1983); Barnwell (1976, 1982); Barnwell and Thurman (1984); Barnwell et al. (1987); Bartell (1967); Bartell and Fingerman (1969); Bartell et al. (1971); Basan and Frey (1977); Becker and Hinsch (1982); Behum et al. (2005); Bergey and Weis (2006); Bergin (1978, 1981); Berglund et al. (1996); Bildstein et al. (1989); Bliss (1982); Bolau (1878); Borst et al. (1987); Bosc (1802); Botos et al. (2000); Bott (1973); Breteler et al. (1981); Brodie and Godley (2006); Brodie and Staton (2003, 2004); Brodie et al. (2005); Brookins and Epifanio (1985); Brown et al. (1954 x 2; 1979); Burkenroad (1947); Burton and Hopkins (2002); Butler and Fingerman (1984, 1986); Callahan and Weis (1983); Callicott and Mangum (1993); Cameron and Forward (1993); Capaldo (1993); Carlson (1935, 1936); Castiglioni and Negreiros-Fransozo (2005, 2006); Chiussi and Díaz (2001, 2002); Christiansen and Yang (1976); Christy (1978, 1979, 1980, 1982, 1983, 1987, 1988 x 2, 1989, 2003); Christy and Backwell (2006); Christy and Salmon (1984, 1991); Christy and Schober (1994); Chrzanowski and Cowley (1977); Chung et al. (1994, 1995, 1997, 1998 x 2); Clark (1896); Clark et al. (1986); Claybrook (1976); Clifford (1982); Colby and Fonseca (1980, 1981, 1984); Coleman (1982); Coohill (1982); Coohill and Fingerman (1975, 1976); Coohill et al. (1969, 1970); Costa et al. (2006); Coventry (1944); Coward et al. (1970); Cowley and Frye (1978); Cowley and Chrzanowski (1979, 1980); Crane (1941, 1943, 1944, 1957, 1958, 1966, 1975); Crane et al. (2004); Cripe and Livingston (1977); Croll and Klinger (1994); Cuiffreda (1994); Cunningham and Myers (1987); Curran and Martin (2003); Currin et al. (1995); Daleo et al. (2003); Dall (1971); Dame and Vernberg (1973, 1978); Davis (1978); Davis and Hagadom (1982); de Araujo et al. (1995); De Kay (1844 = G. vocans); De Vries and Wolcott (1991); De Vries et al. (1991); Dean and Vernberg (1965, 1966); DeCoursey (1979, 1980); DeCoursey and Vernberg (1972); Deecaraman and Fingerman (1985, 1992); Dembowski (1926); Démeusy (1957); Denger and Tankersley (2005); DePatra and Levin (1989); deRivera (2003); deRivera and Vehrencamp (2001); Desmarest (1825); Devi et al. (1993); Dietrich (1979); Dingle (1983); Dircksen et al. (1988); Dittel R. and Epifanio (1982); Doherty (1982); D'Orazio and Holliday (1985); Dores and Herman (1980, 1981); Drews et al. (1982); Dukes et al. (2004); Duncker (1903); Dunn et al. (1990); Durica and Hopkins (1996); Durica et al. (1995, 1997, 1999, 2000, 2001, 2002, 2003, 2005, 2006); Duval (1994, 1995, 1996); Duval and Rittschof (1997); Eastmen-Reks and Fingerman (1984, 1985); Edney (1961); Edwards (1950); Eisen and Jeffrey (1969); Eisen et al. (1973); Engel (1973); Engel et al. (1971); Evans et al. (1976); Ewing (1989); Felder and Staton (1994); Fielder et al. (1971 x 2); Fingerman (1957, 1970, 1973); Fingerman and Bartell (1969, 1973); Fingerman and Couch (1967); Fingerman and Fingerman (1971, 1972, 1974 x 2, 1975, 1976, 1977 x 4, 1978 x 2, 1979 x 2, 1980); Fingerman and Fitzpatrick (1956); Fingerman and Rao (1968, 1969 x 2, 1970); Fingerman and Yamamoto (1964, 1967); Fingerman et al. (1958, 1964, 1966 x 2, 1969, 1971, 1974 x 3, 1978, 1979, 1981 x 2, 1985 x 2, 1994 x 2, 1998); Forward (1977); Forward and Cronin (1979); Forward and Rittschof (1994); Forward et al. (2001 ,2003); Fowler (1912); Frey et al. (1973); Frix et al. (1991); Full and Herreid (1984); Full et al. (1982); Gäde (1979); Gaus et al. (1989); Genoni (1985, 1991); George et al. (2001); Gerald et al. (2005); Gould (1841 = G. vocans); Grant and Eisen (1980); Grant et al. (1980, 1981, 1983); Graszynski and Bigalke (1986); Graszynski and Drews (1981); Graszynski and Rohloff (1983); Graszynski et al. (1979, 1981); Gray (1942); Green et al. (1959); Greenberg et al. (1983); Greenspan (1982); Grimes (1976); Gudmundsdóttir (2002); Gulmann and Mullineaux (2001); Guyselman (1953); von Hagen (1961, 1962, 1970, 1980); Hammerstein (1981); Hanke et al. (1992); Hannan and Evans (1973); Hanström (1935); Hanumante and Fingerman (1981 x 3, 1982 x 7); Hanumante et al. (1980, 1981, 1984); Haresign (1986); Haresign et al. (1996); Hariharan (1966); Hartnoll (1988, 2006); Harwitz et al. (1983); Hay and Shore (1918); Heard (1977); Heit (1974); Heit and Fingerman (1973, 1975); Hemmi (2005); Hemmi and Zeil (2003); Henmi (2003); Herman (1975); Herman and Dallmann (1975); Herreid (1969); Herreid and Mooney (1984); Herrnkind (1966, 1967, 1968 x 3, 1972, 1983); Herrnkind et al. (1968); Honegger (1973, 1976); Hopkins (1981, 1982 x 2, 1983, 1985 x 2, 1986, 1988, 1989 x 3, 1992, 1993, 1994, 1996, 1999, 2001, 2003); Hopkins and Durica (1995); Hopkins and Mislan (1986); Hopkins and Watson (1983); Hopkins et al. (1994, 1997, 1999, 2003); Horch and Salmon (1969); Horch et al. (1980, 2002); Horst (1995); Houser and Allen (1996); Hughes and Seed (1995); Hurman (2005); Huxley and Callow (1933); Hsiang and Premuzic (1975); Hwang and Oswald (1982); Hyatt (1974, 1975, 1977 x 2, 1983); Hyatt and Salmon (1978, 1979); Hyatt et al. (1979); Hyman (1920 x 2); Iribarne and Martinez (1999); Ives (1891); Jaramillo and Lunecke (1988); Jaroensutasinee and Tantichodok (2002); Jaros (1990); Jennions et al. (2003); Jimenez and Bennett (2005); Johnson (1974, 2003); Jones (1968); Jordão and Oliveira (2001, 2005); Jordão et al. (2007); Jorenby et al. (1987); Kanchanaphum et al. (1998); Keller and Andrew (1973); Keller and Wunderer (1978); Kerwin (1971); Kim and Choe (2003); Kim et al. (2004, 2005); Kingsley (1878, 1879, 1880); Kinne (1963); Klaassen and Ens (1993); Klein and Lincer (1974); Klein et al. (1992); Kleinholz (1970, 1972, 1975, 1976); Kleinholz and Keller (1973); Kleinholz and Kimball (1965); Kleinholz et al. (1986); Kneib et al. (1980); Knopf (1966); Ko and Kim (1989); Koch and Gillen (2004); Koga et al. (1998, 2000); Koretsky et al. (2002); Kraeuter (1976); Krebs et al. (1974); Krishnakumaran and Schneiderman (1968); Kulkarni and Fingerman (1985, 1986 x 3, 1987, 1991, 1992); Lambert (1977); Lambert and Crowe (1973, 1976); Lambert and Epifanio (1982); Lambert and Fingerman (1976, 1978); Lambert et al. (1975); Land and Layne (1995 x 2); Landsperger et al. (1981); Langdon (1971); Langdon and Herrnkind (1985); Latreille (1802-1803, 1817); Latruffe et al. (1999); Layne (1998); Layne et al. (1997, 2003 x 2); Le Conte (1855); LeBlanc et al. (2006); Lecroisey and Keil (1985); Lecroisey et al. (1987); Lecroy and Wagner-Merner (1976); Lee et al. (1981, 1982); Leija-Tristán et al. (1990); Leuven et al. (1982); Levinton and Allen (2005); Levinton and Judge (1993); Levinton et al. (1996); Lim (2004, 2006); Lim and Deong (2003); Lim and Heng (2007); Lin et al. (2002); Liñán-Cabello et al. (2002); Litulo (2004, 2005 x 2); Liu and Laufer (1996); López-Duarte and Tankersely (2005); Lucu (1990); Lüschen et al. (1991); Maccagno (1928); Macintosh (1988); Mangum (1993, 1996); Mangum and Greaves (1996); Mangum and Reese (1990); Mantel and Levin (1973); Manwell and Baker (1963); Martin et al. (1984); Masunari (2006); Masunari et al. (2005); Matsumasa and Murai (2005); Mattson (1988); Mayer (1905); McCallum et al. (1991); McConnell and Shelby (1970); McCraith et al. (2003); McGrath et al. (1994); McLain et al. (2003, 2005); McMahon et al. (1990); Menzies and George (1972); Milke and Kennedy (2001); Miller (1961, 1965, 1973); Miller and Maurer (1973); Miller and Vernberg (1968); Milne Edwards (1852); Moffett (1985); Mohrherr and Rao (1985); Mooney and Herreid (1983); Moore (1991); Moore et al. (1968); Morgan (1920); Morgan and Christy (1997); Morrell et al. (2005); Mortin and Marder (1991); Mulstay (1987); Murai and Backwell (2005); Murai et al. (1983, 1987, 2002); Nagabhushanam et al. (1979, 1994); Nakasone et al. (1983); Nässel et al. (1991); Negreiros-Fransozo et al. (2003); Nobbs and McGuinness (1999); Nomann and Pennings (1998); Novak and Salmon (1974); O'Connor (1987, 1990, 1991, 1992, 1993); O'Hara (1973 x 2); Oliveira and Custódio (1998); Oliveira et al. (1998); Olivier (1811); Orr (1955); Ortmann (1897); Palmer (1973; 1988, 1989, 1990 x 3); Pawlik et al. (1980); Payen (1977); Pearse (1914 x 2, 1928); Pelkie (1973); Pennington et al. (2004); Perona et al. (1997); Peters (1955); Petit and Bildstein (1987); Pezalla et al. (1978); Pitts and Cowley (1974); Plumley et al. (1980); Pope (1997, 2000 x 2); Powers (1975, 1977, 1979); Pratt and McLain (2002, 2006); Pratt et al. (2002, 2003, 2005); Quackenbush (1981); Quackenbush and Fingerman (1982, 1983, 1984 x 3, 1985); Quackenbush and Herrnkind (1980, 1981 x 2); Quackenbush and Keeley (1987, 1988); Quackenbush and Rao (1979); Rao (1978); Rao and Fingerman (1968, 1969, 1970 x 2); Rao and Hackett (1973); Rao and Milton (1970); Rao and Riehm (1988 x 2, 1989); Rao et al. (1967 x 2, 1968 x 2, 1970 x 2, 1971, 1972, 1984, 1985, 1986, 1987); Rathbun (1900, 1905, 1918); Rawson and DeCoursey (1976); Rebach (1983); Rebach and French (1996); Reddy and Fingerman (1995); Reddy et al. (1996, 1997); Reinsel (1991, 1992, 1993, 2004); Reinsel and Rittschof (1995); Rheinsmith et al. (1974); Rhodes (1986); Ribeiro et al. (2005); Richardson et al. (1991); Riehm and Rao (1985, 1987); Riehm et al. (1985); Ringold (1979); Rittschof et al. (1995); Robertson(1980); Robertson and Newell (1982 x 2); Robertson and Pfeiffer (1982); Robertson et al. (1980, 1981); Roccatagliata and Jordá (2002); Rodríguez et al. (1992, 2000); Rosenberg (2000, 2001, 2002); Rothe et al. (1991); Rudenskaya (2003); Rudenskaya et al. (2004); Ruppert and Fox (1988); Saigusa et al. (2003); Salmon (1962, 1965, 1966, 1967, 1971, 1983, 1984 x 2, 1987); Salmon and Atsaides (1968 x 2, 1969); Salmon and Horch (1972, 1973); Salmon and Hyatt (1979, 1983 x 2); Salmon and Stout (1962); Salmon and Zucker (1988); Salmon et al. (1977, 1978, 1979); Salomon and Graszynski (1982); Sandeen (1950); Sandifer (1975); Sarojini et al. (1992, 1993, 1995 x 2); Say (1817-1818); Schoor et al. (2000); Schwartz and Safir (1915 x 2); Scott and Mote (1974); Sears and Rittschof (1991); Sears et al. (1991); Sedimeter and Keller (1978); Seed and Hughes (1997); Selander et al. (1971); Semmes et al. (1985); Sendler (1912); Seno and Shigemoto (2007); Severinghaus and Lin (1990); Shih (1999); Shin et al. (2004); Silliman et al. (2003); Silverthorn (1973, 1975 x 2); Smith (1870); Smith and Miller (1973); Spindler et al. (1976); Spindler-Barth (1976); Spivak et al. (1991); Staub (1983); Staub and Fingerman (1982, 1983, 1984 x 2); Stephensen (1921); Stewart and Green (1969); Stimpson (1859); Sturmbauer et al. (1996); Subburajo et al. (1988); Sullivan and Tentori (1981); Sullivan et al. (1983); Summers (1968); Takahashi and Seifter (1972); Takeda and Murai (1993); Takeda and Yamaguchi (1973); Tashian (1956); Tashian and Vernberg (1958); Tautz (1990); Tazelaar (1933); Teal (1958, 1959, 1962); Teal and Carey (1967); Teller (1982, 1984); Teller and Graszynski (1982, 1983); Thurman (1979, 1998, 2002, 2003, 2004 x 2); Thurman and Broghammer (2001); Tilden et al. (1997, 2000, 2001 x 3, 2003 x 2, 2004); Thompson et al. (1989); Thurman (1988, 2001, 2003, 2005); Trinkaus-Randall (1982); Tsai et al. (1991); Tsu and Craik (1996); Tsu et al. (1993, 1994, 1997); Turpin et al. (1970); Vacca (1973); Vacca and Fingerman (1975 x 2, 1983); Vannini (1980); Vernberg and Costlow (1966 x 2); Vernberg and O'Hara (1972); Vernberg and Tashian (1959); Vernberg and Vernberg (1966 x 2, 1969, 1972, 1975, 1983); Vernberg et al. (1974 x 2, 1978); Verrill (1873); Verrill and Smith (1974); Vigh and Fingerman (1982, 1985); Viscido and Wethey (2002); Viscido et al. (2001, 2002); Wada and Murata (2000); Wagner-Merner (1979); Wanson et al. (1984); Watts (1988); Weaver and Salmon (2002); Webb (1977, 1983); Webb and Stokes Lewis (1977); Webb et al. (1954); Weinstein (1993); Weinstein and Full (2000); Weinstein et al. (1985, 1987); Weis (1976 x 3, 1977 x 2, 1978 x 2, 1980, 1985); Weis and Kim (1986, 1988); Weis and Mantel (1976); Weis and Perlmutter (1987 x 2); Weis and Weis (1979, 1988, 1992, 2004); Weis et al. (1986, 1987 x 2, 1991, 1992, 2002); Weissburg (1991, 1992, 2001); Weissburg and Zimmer-Faust (1991); Weissburg et al. (1996, 2001); Welgus and Grant (1983); Welgus et al. (1982); Wilkens (1960); Wilkens and Fingerman (1965); Windsor et al. (2005); Winger et al. (1990); Winstead and Thurman (2006); Wlodawer et al. (2004); Wolin and Laufer (1971); Wolin et al. (1973); Wong et al. (2005); Wright and Roosenburg (1982); Wright et al. (1984); Wu et al. (1999, 2002, 2003, 2004 x 2); Yamaguchi (1971, 1977, 2001 x 5, 2002, 2003); Yamaguchi and Henmi (2001); Yamaguchi and Tabata (2004); Yamaguchi et al. (2005 x 2); Yerkes (1901); Yoder et al. (2005); Young (1900); Young and Ambrose (1978); Young and Govind (1983); Zahnow and Rao (1985); Zeil and Al-Mutairi (1996); Zeil and Layne (2002); Zeil et al. (1986); Zeleny (1905); Zhong et al. (2003); Zimmer et al. (1999); Zou (2003, 2005 x 2); Zou and Bonvillain (2003 x 2, 2004 x 2); Zou and Fingerman (1998, 1999 x 5, 2000); Zucker (1978, 1983, 1984)